Chapter Concepts 1.1 The Scientic Process The scientic process allows biologists to gather information and come to conclusions about the natural world. 2 Various conclusions pertaining to the same area of interest are sometimes used to arrive at a theory, a general concept about the natural world. 5 All persons have the responsibility to decide how scientic information can best be used to make ethical or moral decisions. 6 1.2 The Characteristics of Life Although life is quite diverse, it can be dened by certain common characteristics. 7 The biosphere is made up of ecosystems where living things interact with each other and the physical environment. 10 The biodiversity of the biosphere is being threatened by human activities. 12 1.3 The Classication of Living Things Living things are classied into categories according to their evolutionary relationships. 13 Many biological experiments are performed in the laboratory where conditions can be more easily controlled. Back Forward Main Menu TOC Study Guide TOC Textbook Website Student OLC MHHE Website 1.1 The Scientic Process Biology is the scientic study of life (Fig. 1.1). Religion, aes- thetics, ethics, and, yes, science, are all ways that human beings have of nding order in the natural world. But sci- ence differs from other human ways of knowing and learn- ing by its process. The process of science is unique. Science is a human endeavor that considers only what is observable by the sens- 2 Introduction 1-2 A bout 200 years ago, an English country doctor, Edward Jenner, made medical history due to a keen observation. He noticed that milkmaids usually avoid- ed contracting smallpox, a disguring and frequently fatal disease. Jenner reasoned that the women were somehow protected because they had previously had cowpox, a simi- lar but much less serious illness. To test that hypothesis, Jenner injected a young boy with cowpox, and later exposed him to smallpox. When his volunteer remained healthy, the way was paved for the development of the mod- ern version of a smallpox vaccine that has done away with the disease entirely. Biology uses the same methodology today as was used by Jenner to come to conclusions that are accepted until proven false by future investigations. The information pre- sented in this text is the result of work done by scientists over the ages. What is science and what methodology does science use? Figure 1.1 Diversity of life. Biology is the scientic study of life, which has diversied into the many forms we nd on planet Earth. a. Vorticella, a protozoan; b. Hibiscus, a owering plant; c. Sally lightfoot, a crab; d. Snow leopard, a mammal es or by instruments that extend the ability of the senses. Microscopes extend the ability of sight way beyond what can be seen by the naked eye, for example. Observations can be made by any one of our ve senses; we can observe with our noses that dinner is almost ready; observe with our n- gertips that a surface is smooth and cold; and observe with our ears that a piano needs tuning. Various input allows sci- entists to formulate hypotheses. Formulating a Hypothesis Once a scientist becomes interested in an observable event, called a phenomenon, he or she will most likely study up on it. The goal will be to nd any other past work in the area; the Internet, the library, other scientists will all be consulted to see what past ideas there may have been about this phe- nomenon. All of this input helps a scientist use imagination and creative thinking to formulate a hypothesis. Ahypothe- sis is a tentative explanation of a scientic question based on observations and past knowledge. Hypotheses, of course, vary in the level of their sophis- tication. A young person in elementary school might hypothesize that plants need light in order to grow; while a high school student might hypothesize that light allows plant leaves to produce sugar. In any case, all the past expe- riences of the individual no matter their source, will most likely inuence the formulation of a hypothesis. Einstein said that aesthetic qualities like beauty and simplicity 30 m d. Snow Leopard, a mammal a. Vorticella, a protozoan b. Hibiscus, a flowering plant c. Sally lightfoot, a crab Back Forward Main Menu TOC Study Guide TOC Textbook Website Student OLC MHHE Website helped him to formulate his hypotheses about the workings of the universe. Indeed, most every scientist looks favorably upon the simplest hypothesis for a phenomenon and tests that one rst. Science only considers hypotheses that can be tested. Moral and religious beliefs while very important to our lives differ between cultures and through time and they are not always testable by further observations and/or experimen- tation. Testing a Hypothesis Hypotheses are tested either in the laboratory setting or in a natural setting. The laboratory is a place that lends itself to carrying on experiments, which are articial situations devised to test hypotheses. Experiments can also be car- ried out in a natural setting, called the field. David P. Barash, who was observing the mating behavior of moun- tain bluebirds, decided to perform an experiment to test the hypothesis that aggression of the male varies accord- ing to the reproductive cycle. He posted a male bluebird model near nests while the resident male was out forag- ing. The behavior of the male toward the model and toward his female mate were noted during the first ten minutes of the males return. Aggression was most severe when the male model was presented before the rst egg was laid, less severe when the model was presented after an egg was laid, and least severe after the eggs had hatched (Fig. 1.2). Laboratory experiments are often preferred to experi- ments in the eld because a laboratory setting eliminates extraneous variables. Avariable is a factor that can cause an observable change during the progress of the experiment. The Science Focus on the next page describes a hypothetical experiment scientists conduct to determine whether sub- stance S is a safe food additive. Some of the possible vari- ables include the individual characteristics of the mice, availability of food and water, and environmental condi- tions such as the cleanliness of the cages and the tempera- ture and humidity of the room, and so forth. The experimenters would try to keep all these variables constant except the presence or absence of substance Experiments are considered more rigorous when they include a control group. Control in this context does not refer to the power of the experimenterrather it refers to the validity of the experimental results. A control group experiences all the steps in the experiment except for the one that is being tested. Barash used a control group. He posted a male robin instead of a male bluebird near certain nests. (The male bluebirds did not respond at all to a robin model.) In the substance S experiment, the mice in the con- trol group do not have substance S added to their food. When an experiment has a control group, we know that the results are due to the variable being tested and not due to some nonidentiable chance event that occurred during the experiment. Chapter 1 The Study of Life 3 1-3 Stage of Nesting Cycle A p p r o a c h e s
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M i n u t e nest construction 0.5 1.0 1.5 2.0 approaches to male model approaches to female bird nest 1 nest 2 key: 0 first egg laid hatching of eggs c. Scientist studies results and comes to a conclusion. b. Scientist performs experiment and collects objective data. a. Scientist makes observations, studies previous data, and formulates a hypothesis. Figure 1.2 Example of scientic method. Observation of male bluebird behavior allowed David Barash to formulate a testable hypothesis. Then, he collected the data displayed in a graph and came to a conclusion. Back Forward Main Menu TOC Study Guide TOC Textbook Website Student OLC MHHE Website Biologists want to determine if sweetener S is a safe food addi- tive and design a controlled laboratory experiment. They place a certain number of randomly chosen inbred (genetically simi- lar) mice into the two groupssay, 100 mice per group (Fig. 1A). If any of the mice are different from the others, it is hoped random selection will distribute them evenly among the groups. The researchers also make sure that all conditions, such as availability of water, cage setup, and temperature of the sur- roundings, are the same for both groups. The food for each group is exactly the same except for the amount of sweetener S. In experiment 1, the researchers formulate a hypothesis that sweetener S is a safe food additive at any dietary intake. The control group receives no sweetener S and in the test group 50% of the diet is sweetener S. At the end of the experiment, both groups of mice are examined for bladder cancer. They nd that one-third of the mice in the test group have bladder cancer, while none in the control group have bladder cancer. The results of this experiment falsify the hypothesis and the researchers conclude that sweetener S is not a safe food additive at 50% of dietary intake. The researchers decide to rene their study, and in experiment 2 they hypothesize that sweetener S is safe if the diet contains a limited amount of sweetener S. They feed sweetener S to groups of mice at ever-greater concentrations until it reaches 50% of the diet. Group 1: diet contains no sweetener S (the control) Group 2: 5% of diet is sweetener S Group 3: 10% of diet is sweetener S Z Group 11: 50% of diet is sweetener S The researchers present their data in the form of a graph and statistically analyze their data to determine if the difference in the number of cases of bladder cancer between the various groups is signicant and not due to simple chance. Finding that they are signicant, the researchers conclude that they can now develop a recommendation concerning the intake of sweetener S in humans. They suggest that an intake of sweetener S up to 10% of the diet is relatively safe but thereafter an ever-greater incidence of bladder cancer should be expected (Fig. 1A). A Controlled Laboratory Experiment Figure 1A Design of a controlled experiment. Genetically similar mice are randomly divided into a control group and (a) test group(s) that contain 100 mice each. All groups are exposed to same conditions, such as cage setup, temperature, and water supply. The control group is not subjected to sweetener S in the food. At the end of the experiment, all mice are examined for bladder cancer. Test group: sweetener in food Control group: no sweetener in food Bladder cancer examination Experiment 1 control group: no bladder cancer test group: one-third of mice have bladder cancer Sweetener S in food (%) M i c e
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( % ) 10 15 20 25 30 0 10 20 30 40 50 Experiment 2 Back Forward Main Menu TOC Study Guide TOC Textbook Website Student OLC MHHE Website The Experimental Results The results of an experiment are referred to as the data. Data often takes the form of a table or a graph that allows one to see the results in an organized manner. Barashs data are shown in Figure 1.2. Data should be objective rather than subjective, and mathematical data are conducive to objectiv- ity. For example, Barash didnt report that it seemed to him that the male bluebirds were more aggressive before the eggs were laid (subjective); rather, he dened aggression as the number of approaches per minute (objective) and reported that number. When Mendel, the father of genetics, did a pea cross concerning height, he didnt report that he could tell at a glance there were more tall than short plants; he reported that the ratio was three tall to every short plant. In fact, Mendel repeated one particular experiment so many times that he counted 7,324 peas! To ensure that the results are not due to chance or some unknown variable, experiments are often repeated not only by the original scientist but by others in the same area of expertise. Therefore, scientists must keep careful records of how they perform their experiments so that others can repeat them. If the same results are not obtained over and over again, the experiment is not considered valid. The Conclusion After studying the results, the experimenter comes to a con- clusion whether the results support or falsify (show to be untrue) the hypothesis. The data allowed Barash to con- clude that aggression in male bluebirds is related to their reproductive cycle. Therefore, his hypothesis was support- ed. If male bluebirds were always aggressive even toward male robin models, his hypothesis would have been proven false. Science progresses and the hypothesis may have to be modied in the future. There is always the possibility that a more sophisticated experiment using perhaps more advanced technology might falsify the hypothesis. There- fore, a scientist never says that the data prove the hypoth- esis to be true. Because of this feature, some think of science as what is left after alternative hypotheses have been rejected. Scientists report their ndings in scientic journals so that their methodology and results are available to the scien- tic community. Barash reported his experiment in the American Naturalist. 1 The reporting of experiments results in accumulated data that will help other scientists formulate hypotheses. Also, it results in a body of information that is made known to the general public through the publishing of books, such as this biology textbook. Scientic Theories The ultimate goal of science is to understand the natural world in terms of scientic theories, which are concepts that join together well-supported and related hypotheses. In a movie, a detective may claim to have a theory about the crime. Or you may say that you have a theory about the won-lost record of your favorite baseball team. But in sci- ence, the word theory is reserved for a conceptual scheme that is supported by a broad range of observations, experi- ments, and data. Some of the basic theories of biology are: Name of Theory Explanation Cell All organisms are composed of cells. Biogenesis Life comes only from life. Evolution All living things have a common ancestor and are adapted to a particular way of life. Gene Organisms contain coded information that dictates their form, function, and behavior. Evolution is the unifying concept of biology because it per- tains to various aspects of living things. For example, the theory of evolution enables scientists to understand the his- tory of life, the variety of living things, and the anatomy, physiology, and development of organismseven their behavior. Barash gave an evolutionary interpretation to his results. It was adaptive, he said, for male bluebirds to be less aggressive after the rst egg is laid because by that time the male bird is sure the offspring is his own and maladaptive for the male bird to waste time and energy being too aggres- sive toward a rival and his mate after hatching because his offspring is already present. (When an organism is adapted to its environment, it is better able to survive and produce offspring.) Fruitful theories are ones that help scientists generate new hypotheses, and the theory of evolution has been a very fruitful theory. In fact, it probably helped Barash develop the hypothesis he chose to test. Because the the- ory of evolution has been supported by so many obser- vations and experiments for over 100 years, some biologists refer to the principle of evolution, suggesting that this is the appropriate term for theories that are gen- erally accepted by an overwhelming number of scien- tists. The term law instead of principle is preferred by some. In another chapter concerning energy relation- ships, for example, we will examine the laws of thermo- dynamics. Chapter 1 The Study of Life 5 1-5 1 Barash, D. P. 1976. The male responds to apparent female adultery in the mountain bluebird, Sialia currucoides: An evolutionary interpretation. American Naturalist 110:1097101. Back Forward Main Menu TOC Study Guide TOC Textbook Website Student OLC MHHE Website The Scientic Method The process of science is often described in terms of the sci- entic method. Some scientists object to outlining the steps of the scientic method as is done in Figure 1.3 because such a diagram suggests a rigid methodology. Actually, sci- entists approach their work in many different ways and even sometimes make discoveries by chance. The most famous case pertains to penicillin. When examining a petri dish in 1928, Alexander Fleming noticed an area around a mold that was free of bacteria. Upon investigating, Fleming found that the mold produced an antibacterial substance he called penicillin. Penicillin was later mass produced and is still a successfully used antibiotic in humans today. Some scientists do not test hypothesesinstead, they make observations and add to the known data. For example, a sci- entist might decide to nd out what types of animals live on the ocean oor. A discussion of the scientic process is not complete without an admission that scientists do have to accept cer- tain assumptions. They have to believe, for example, that nature is real and understandable and knowable by observ- ing it; that nature is orderly and uniform; that measure- ments yield knowledge of the thing measured; and that natural laws are not affected by time. Science and Social Responsibility Science seeks only a natural cause for the origin and history of life. Doctrines of creation that have a mythical, philosoph- ical, or theological basis are outside the realm of science because they cannot be tested by observation and/or exper- imentation. Creationism, which states that God created all species as they are today, cannot be considered science because explanations based on supernatural rather than nat- ural causes involve faith rather than data. There are many ways in which science has improved our lives. The discovery of antibiotics and vaccines has expanded the human life span. Cell biology research is helping us understand the causes of cancer. Genetic research has produced new strains of agricultural plants that have eased the burden of feeding our burgeoning world population. Still there are other instances in which science has resulted in technologies that have harmed the environment. Technology is a process, an instrument, or a structure that is developed or constructed using scientic principles. Biochemical knowledge was used to develop pesticides which have helped increase agricultural yields. Pesticides, as you may know, kill not only pests but also other types of organisms. The book Silent Spring was writ- ten to make the public aware of the harmful environmen- tal effects of pesticide use. Too often we blame science for these developments and think that scientists are duty bound to pursue only those avenues of research that are consistent with a certain system of values. But making value judgments is not a part of science. Ethical and moral decisions must be made by all people. The responsibility for how we use the fruits of science, including a given technology, must reside with people from all walks of life, not upon scientists alone. Sci- entists should provide the public with as much informa- tion as possible when such issues as the use of atomic energy, fetal research, and genetic engineering are being debated. Then they, along with other citizens, can help make decisions about the future role of these technologies in our society. All men and women have a responsibility to decide how to use scientic knowledge so that it benets the human species and all living things. 6 Introduction 1-6 previous data formulation of hypothesis observations and/or experimentation new data conclusion theory 1 2 3 observations Figure 1.3 Flow diagram for the scientic method. On the basis of observations and previous data, a scientist formulates a hypothesis. The hypothesis is tested by further observations or a controlled experiment, and new data either support or falsify the hypothesis. The return arrow indicates that a scientist often chooses to retest the same hypothesis or to test a related hypothesis. Conclusions from many different but related experiments may lead to the development of a scientic theory. For example, studies in biology of development, anatomy, and fossil remains all support the theory of evolution. Back Forward Main Menu TOC Study Guide TOC Textbook Website Student OLC MHHE Website 1.2 The Characteristics of Life Biological theories help us determine the characteristics of life. Despite its diversity (see Figure 1.1), certain properties characterize all living things. For example, all living things are organized. Living Things Are Organized Living things have levels of organization from the atoms that constitute all matter to an ecosystem in which they live (Fig. 1.4). Atoms join together to form molecules such as the DNA molecules that occur only within cells. The cell is the lowest level of biological organization to have the characteristics of life. Anerve cell is one of the types of cells in the body of a white-tailed deer. Atissue is a group of similar cells that perform a particular function. Nervous tissue has millions of nerve cells that transmit signals to all parts of the deers body. Several tissues join together to form an organ. The main organs in the nervous system of a deer are the brain, the spinal cord, and the nerves. Organs work together to form an organ system. The brain sends messages to the spinal cord which, in turn, sends them to body parts by way of the spinal nerves. An organ- ism, meaning an individual living thing, is a collection of organ systems. Adeer contains a digestive system, a circu- latory system, and several other systems in addition to a nervous system. Organisms usually live within a popula- tion, which is a group of interbreeding organisms in a par- ticular locale. Several different populations interact within a community. For example, deer feed on many different types of living plants. An ecosystem includes a communi- ty and also the physical environment. Organisms not only interact with one another, they also interact with the phys- ical environment, as when a deer takes a drink of water from a pond. Chapter 1 The Study of Life 7 1-7 molecular level (DNA) cellular level (nerve cell) tissue level (nervous tissue) organ level (brain) organ system level (nervous system) organism level (white-tailed deer) other levels (population, community, ecosystem) Figure 1.4 Levels of biological organization. Back Forward Main Menu TOC Study Guide TOC Textbook Website Student OLC MHHE Website 8 Introduction 1-8 Living Things Acquire Materials and Energy Living things cannot maintain their organization nor carry on lifes other activities without an outside source of materi- als and energy. Photosynthesizers, such as trees, use carbon dioxide, water, and solar energy to make their own food. Human beings and other animals, like ospreys (Fig. 1.5), acquire materials and energy when they eat food. Food provides nutrient molecules, which are used as building blocks or for energy. Energy is the capacity to do work, and it takes work to maintain the organization of the cell and of the organism. When nutrient molecules are used to make their parts and products, cells carry out a sequence of synthetic chemical reactions. Some nutrient molecules are broken down completely to provide the necessary energy to carry out synthetic reactions. Most living things can convert energy into motion. Self- directed movement, as when we decide to rise from a chair, is even considered by some to be a characteristic of life. Living Things are Homeostatic Homeostasis means staying the same. Actually, the inter- nal environment stays relatively constant; for example, the human body temperature uctuates slightly during the day. Also, the bodys ability to maintain a normal internal tem- perature is somewhat dependent on the external tempera- turewe will die if the external temperature becomes overly hot or cold. All human systems contribute to homeostasis. The digestive system provides nutrient molecules; the circula- tory system transports them about the body; and the excre- tory system rids blood of metabolic wastes. The nervous and hormonal systems coordinate the activities of the oth- er systems. One of the major purposes of this text is to show how all the systems of the human body help to main- tain homeostasis. Living Things Respond to Stimuli Living things respond to external stimuli, often by moving toward or away from the stimulus. Movement in humans is dependent upon their nervous and muscular systems. Other living things use a variety of mechanisms in order to move. Leaves of plants track the passage of the sun during the day, and when a houseplant is placed near a window, its stem bends to face the sun. The movement of an organism, whether self-directed or in response to a stimulus, constitutes a large part of an organisms behavior. Behavior largely is directed toward minimizing injury, acquiring food, and reproducing. Figure 1.5 Living things acquire materials and energy. An osprey is a large bird that preys on shes as its source of food. Figure 1.6 Living things reproduce. An osprey lays two to four eggs in a large nest located on the top of a tree, a rock pinnacle, or even a telephone pole. The edglings are ready to leave the nest 40 to 50 days after hatching. Back Forward Main Menu TOC Study Guide TOC Textbook Website Student OLC MHHE Website Chapter 1 The Study of Life 9 1-9 Living Things Reproduce Life comes only from life. The presence of genes in the form of DNA molecules allows cells and organisms to reproduce that is, make more of themselves (Fig. 1.6). DNAcontains the hereditary information that directs the structure and metabo- lism, chemical reactions of a cell. Before reproduction occurs, genes are replicated and copies of genes are produced. Unicellular organisms reproduce asexually simply by dividing. The new cells have the same genes and structure as the single parent. Multicellular organisms usually repro- duce sexually. Each parent, male and female, contributes roughly one-half the total number of genes to the offspring, which then does not resemble either parent exactly. Living Things Grow and Develop Growth, recognized by an increase in size and often the num- ber of cells, is a part of development. In humans, develop- ment includes all the changes that take place between conception and death. First, the fertilized egg develops into a newborn, and then a human goes through the stages of child- hood, adolescence, adulthood, and aging. Development also includes the repair that takes place following an injury. All organisms undergo development. Figure 1.7 illus- trates that an oak tree progresses from an acorn to a seedling before it becomes an adult oak tree. Living Things Are Adapted Adaptations are modifications that make an organism suited to its way of life. Consider, for example, a bird like an osprey (see Fig. 1.5), which catches and eats sh. An osprey can fly in part because it has hollow bones to reduce its weight and ight muscles to depress and ele- vate the wings. When an osprey dives, its strong feet take the rst shock of the water and then its long and sharp claws hold onto its slippery prey. Adaptations come about through evolution. Evolu- tion is the process by which characteristics of species (a group of similarly constructed organisms that successfully interbreed) change through time. When new variations arise that allow certain members of the species to capture more resources, these members tend to survive and to have more offspring than the other unchanged members. Therefore, each successive generation will include more members with the new variation. In the end, most mem- bers of a species have the same adaptations to their envi- ronment. Evolution, which has been going on since the origin of life, explains both the unity and the diversity of life. All organisms share the same characteristics of life because their ancestry can be traced to the rst cell or cells. Organ- isms are diverse because they are adapted to different ways of life. need confirmation of photo Figure 1.7 Living things grow and develop. Stages in the development of an oak tree from an acorn to a seedling to an adult. Back Forward Main Menu TOC Study Guide TOC Textbook Website Student OLC MHHE Website Living Things Belong to a Population Individual organisms belong to a population, all the mem- bers of a species that live within a particular community. The populations within a community interact among them- selves and with the physical environment (soil, atmosphere, etc.), thereby forming an ecosystem. All ecosystems are included within the biosphere, a thin layer of life that encir- cles the earth. Although an ecosystem like a tropical rain forest changessome trees fall and some animals diean ecosys- tem remains recognizable year after year. We say it is in dynamic balance. In many cases, even the extinction of species (and their replacement by new species through evo- lution) still allows the dynamic balance of the system to be maintained. Amajor feature of the interactions between populations pertains to who eats whom. Plants produce food, and ani- mals that eat plants are food for other animals. Such a sequence of organisms is called a food chain (Fig. 1.8). Both plants and animals interact with the physical environment, as when they exchange gases with the atmosphere. Nutrients cycle within and between ecosystems. Plants take in inorganic nutrients, like carbon dioxide and water, and produce organic nutrients, such as carbohydrates, that are used by themselves and various levels of animal con- sumers. When these organisms die and decay, inorganic nutrients are made available to plants once more. The blue arrows in Figure 1.8 show how chemicals cycle through the various populations of an ecosystem. In contrast, the yellow arrows in Figure 1.8 show how energy ows through an ecosystem: solar energy used by plants to produce organic food is eventually converted to heat when organisms, including plants, use organic food as an energy source. Therefore, a constant supply of solar ener- gy is required for an ecosystem and for life to exist. All living things share the same characteristics of life: they are organized; take materials and energy from the environment; are homeostatic; respond to stimuli; reproduce; grow and develop; are adapted to their way of life; and belong to populations. 10 Introduction 1-10 death and decomposition Key: energy nutrients heat inorganic nutrients heat heat heat heat Figure 1.8 Ecosystem organization. Within an ecosystem, nutrients cycle (see blue arrows); plants make and use their own organic food, and this becomes food for several levels of animal consumers, including humans. When these organisms die and decompose, the inorganic remains are used by plants as they produce organic food. Energy ows (see yellow arrows); solar energy used by plants to produce organic food is eventually converted to heat by all members of an ecosystem (therefore, a constant supply of solar energy is required for life to exist). Back Forward Main Menu TOC Study Guide TOC Textbook Website Student OLC MHHE Website Tropical Rain Forest, a Terrestrial Ecosystem Tropical rain forests are the most complex ecosystems in the world. They are found near the equator where there is plen- tiful sun and rainfall the entire year. Major rain forests are located in South America (Fig. 1.9), central and west Africa, and Southeast Asia. Rain forests have a multilayered canopy which consists of broad-leaved evergreen trees of different heights. Most animal populations live in the canopy where they interact with each other. Brightly colored birds, such as toucans and macaws, y around eating fruits, buds, and pollen. Other birds, such as long-billed hum- mingbirds feed on nectar often taken from small plants that grow independently on the trees. Tree sloths and spider monkeys are mammals that live in the canopy and are preyed upon by jaguars. Other canopy animals include but- teries, tree frogs, and dart-poison frogs. Many canopy ani- mals, such as bats, are active only at night. Snakes, spiders, and ants are animals that live on or near the ground and not in the canopy. Chapter 1 The Study of Life 11 1-11 Figure 1.9 Tropical rain forest populations interact in the manner described in Fig. 1.8. toucan morpho butterfly jaguar epiphytic orchid dart-poison frog Back Forward Main Menu TOC Study Guide TOC Textbook Website Student OLC MHHE Website So far, nearly 2 million species of organisms have been discov- ered and named. Two-thirds of the plant species, 90% of the nonhuman primates, 40% of birds of prey, and 90% of the insects live in the tropics. Many more species of organisms (perhaps as many as 30 million) are estimated to live in the tropical rain forests but have not yet been discovered. Tropical forests span the planet on both sides of the equator and cover 67% of the total land surface of the earthan area roughly equivalent to our contiguous forty-eight states. Every year humans destroy an area of forest equivalent to the size of Oklahoma. At this rate, these forests and the species they con- tain will disappear completely in just a few more decades. Even if the forest areas now legally protected survive, 5672% of all tropical forest species would still be lost. The loss of tropical rain forests results from an interplay of social, economic, and political pressures. Many people already live in the forest, and as their numbers increase, more of the land is cleared for farming. People move to the forests because inter- nationally nanced projects build roads and open the forests up for exploitation. Small-scale farming accounts for about 60% of tropical deforestation, and this is followed by commercial log- ging, cattle ranching, and mining. International demand for tim- ber promotes destructive logging of rain forests in Southeast Asia and South America. Amarket for low-grade beef encourages the conversion of tropical rain forests to pastures for cattle. The lure of gold draws miners to rain forests in Costa Rica and Brazil. The destruction of tropical rain forests produces only short- term benets but is expected to cause long-term problems. The forests soak up rainfall during the wet season and release it dur- ing the dry season. Without them, a regional yearly regime of ooding followed by drought is expected to destroy property and reduce agricultural harvests. Worldwide, there could be changes in climate that would affect the entire human race. On the other hand, the preservation of tropical rain forests offers benets. For example, the rich diversity of plants and animals would continue to exist for scientic and pharmacological study. One-fourth of the medicines we currently use come from tropical rain forests. The rosy periwinkle from Madagascar has produced two potent drugs for use against Hodgkin disease, leukemia, and other blood cancers. It is hoped that many of the still- unknown plants will provide medicines for other human ills. Studies show that if the forests were used as a sustainable source of nonwood products, such as nuts, fruits, and latex rub- ber, they would generate as much or more revenue while con- tinuing to perform their various ecological functions. And biodiversity could still be preserved. Brazil is exploring the con- cept of extractive reserves, in which plant and animal prod- ucts are harvested but the forest itself is not cleared. Ecologists have also proposed forest farming systems, which mimic the natural forest as much as possible while providing abundant yields. But for such plans to work maximally, the human popu- lation size and the resource consumption per person must be stabilized. Preserving tropical rain forests is a wise investment. Such action promotes the survival of most of the worlds species indeed, the human species, too. 12 Tropical Rain Forests: Can We Live Without Them? The Human Population The human population tends to modify existing ecosystems for its own purposes. As more and more ecosystems are con- verted to towns and cities, fewer of the natural cycles are able to function adequately to sustain an evergrowing human population. It is important to do all we can to pre- serve ecosystems, because only then can we be assured that we will continue to exist. The recognition that the workings of the ecosystems need to be preserved is one of the most important developments of our new ecological awareness. Presently, there is great concern about preserving the worlds tropical rain forests, as discussed in the reading on this page. The tropical rain forests perform many services for us. For example, they act like a giant sponge and absorb carbon dioxide, a pollutant that pours into the atmosphere from the burning of fossil fuels such as oil and coal. If the rain forests continue to be depleted as they are now, an increased amount of carbon dioxide in the atmosphere is expected to cause an increase in the average daily tempera- ture. Problems with acid rain are also expected to increase, since carbon dioxide combines with water to form carbonic acid, a component of acid rain. The present biodiversity (number and size of populations in a community) of our planet is being threatened. It has been estimated that the number of species in the biosphere may be as high as 80 million species, but thus far fewer than 2 million have been identied and named. Even so we may be present- ly losing from 24 to even 100 species a day due to human activities. The existence of the species featured in Figure 1.9 is threatened because tropical rain forests are being reduced in size. Most biologists are alarmed over the present rate of extinction and believe the rate may eventually rival the mass extinctions that have occurred during our planets history. The human population tends to modify existing ecosystems and to reduce biodiversity. Because all living things are dependent upon the normal functioning of the biosphere, ecosystems should be preserved. Back Forward Main Menu TOC Study Guide TOC Textbook Website Student OLC MHHE Website 1.3 The Classication of Living Things Taxonomy is that part of biology dedicated to naming, describing, and classifying organisms. Taxonomists use spe- cic criteria to classify species, including human beings, into certain categories. As we move from genus to kingdom, more and more different types of species are included in each successive cat- share very similar characteristics, but those that are in the same kingdom have only general characteristics in common. In the same way, all species in the genus Zea look pretty much the samethat is, like corn plantswhile species in the plant kingdom can be quite different, as is evident when we compare grasses to trees. Taxonomists give each species a scientic name in Latin. The scientic name is a binomial (bi means two; nomen means name). For example, the name for humans is Homo sapiens, and for corn, it is Zea mays. The rst word is the genus, and the second word is a specic epithet for that species. (Note that both words are in italic but only the genus is capitalized.) Scientic names are universally used by biol- ogists so as to avoid confusion. Common names tend to overlap and often are in the language of a particular country. Taxonomy makes sense out of the bewildering variety of life on earth. Species are classied according to their pre- sumed evolutionary relationship; those placed in the same genus are the most closely related, and those placed in sepa- rate kingdoms are the most distantly related. As more is known about evolutionary relationships between species, taxonomy changes. Presently many biologists recognize the ve kingdoms listed in Figure 1.10. Others disagree not only about the number of kingdoms but also about which species should be placed in the various categories of classication. Chapter 1 The Study of Life 13 1-13 Categories For Humans Description Kingdom Animalia Multicellular, moves, ingests food Phylum Chordata Dorsal supporting rod and nerve cord Class Mammalia Hair, mammary glands Order Primates Adapted to climb trees Family Hominidae Adapted to walk erect Genus Homo Large brain, tool use Species H. sapiens Kingdoms of Life Representative Organisms Organization Type of Nutrition Representative Organisms Monera Protista Fungi Plantae Animalia Microscopic single cell (sometimes chains or mats) Complex single cell, some multicellular Some unicellular, most multicellular filamentous forms with specialized complex cells Multicellular form with specialized complex cells Multicellular form with specialized complex cells Absorb food (some photo- synthesize) Absorb, photosynthesize, or ingest food Absorb food Photosynthesize food Ingest food Bacteria including cyanobacteria Protozoans, algae, water molds, and slime mold Molds, yeast, and mushrooms Mosses, ferns, nonwoody and woody flowering plants Invertebrates, fishes, reptiles, amphibians, birds, and mammals bacilli spirochete Anabaena Gloeocapsa black bread mold yeast mushroom bracket fungus coral earthworm blue jay squirrel moss fern pine tree nonwoody flowering plant paramecium euglenoid slime mold dinoflagellate egory. Only human beings are in the genus Homo, but many different types of animals are in the animal kingdom. Notice that in the example given, species within the same genus Figure 1.10 Classication of organisms. In this text, organisms are classied into the ve kingdoms illustrated in this table. Back Forward Main Menu TOC Study Guide TOC Textbook Website Student OLC MHHE Website Summarizing the Concepts 1.1 The Scientic Process When studying the world of living things, biologists and other scien- tists use the scientic process. Observations along with previous data are used to formulate a hypothesis. New observations and/or experi- ments are carried out in order to test the hypothesis. Scientists often do controlled experiments. The control sample does not go through the step being tested, and this acts as a safeguard against a wrong conclu- sion. The new data may support a hypothesis or they may prove it false. Hypotheses cannot be proven true. Several conclusions in a particular area may allow scientists to arrive at a theorygeneralizations such as the cell theory, gene theory, or the theory of evolution. Evolution is the unifying theory of biology. Science is objective and uses conclusions based on data to arrive at theories about the natural world. Any explanation based on supernat- ural beliefs cannot be considered science because such beliefs are not tested in the usual scientic way. Science does not answer ethical questions; we must do this for ourselves. Knowledge provided by science, such as the contents of this text, can assist us in making decisions that will be benecial to human beings and to other living things. 1.2 The Characteristics of Life Evolution accounts for both the diversity and the unity of life we see about usall organisms share the same characteristics of life: 1. Living things are organized. The levels of biological organization extend from the cell to ecosystems: atoms and molecules cells tissues organs organ systems organisms populations communities ecosystems. In an ecosystem, populations interact with one another and the physical environment. 2. Living things take materials and energy from the environment; they need an outside source of nutrients. 3. Living things are homeostatic; internally they stay just about the same despite changes in the external environment. 4. Living things respond to stimuli; they react to internal and external events. 5. Living things reproduce; they produce offspring that resemble themselves. 6. Living things grow and develop; during their lives they changemost multicellular organisms undergo various stages from fertilization to death. 7. Living things are adapted; they have modications that make them suited to a particular way of life. 8. All living things belong to a population, which can be dened as all the members of the same species that occur in a particular locale. 1.3 The Classication of Living Things Living things are classied according to their evolutionary relation- ships into these ever more specic categories: kingdom, phylum, class, order, family, genus, species. Organisms in different kingdoms are only distantly related; organisms in the same genus are very closely related. Studying the Concepts 1. What are the steps of the scientic process? Why cant this process prove a hypothesis true? 2 2. What is a controlled experiment? Why must a scientist test one variable at a time? have a control group? 3 3. What is the ultimate goal of science? Give an example that supports your answer. 5 4. Why isnt creationism considered a part of science by biolo- gists? 6 14 Introduction 1-14 T he Endangered Species Act requires the federal government to identify endangered and threatened species and to protect their habitats, even to the extent of purchasing their habitats. Developers feel that the act protects wildlife at the expense of jobs for US citizens. In an effort to allow development in sensitive areas, it is now possible to move forward after a Habitat Conservation Plan (HCP) is approved. An HCP permits, say, new-home construction or logging on a part of the land if wildlife habitat is conserved on another part. Con- servation can also mean helping the gov- ernment buy habitat some place else. The nations rst HCP was approved in 1980. It permitted housing construction on San Bruno Mountain near San Francisco, if 97% of the habitat for the endangered mission blue buttery was preserved. That sounds pretty good, but by this time hun- dreds of HCPs have been approved, and the conservation requirement may have slipped a bit. Tim Cullinan, a director of the National Audubon Society, recently found that logging companies in the Pacif- ic Northwest are proposing the exchange of habitat on public land for the right to log privately owned old forests. In other words, nothing has been given up. By now, there are so many HCPs in the works they are being rubber-stamped by govern- ment ofcials with no public review at all. Do you favor development over preservation of habitat or vice versa? Do you think the federal government should be in the business of trying to preserve endangered species? Do you think that public review of HCPs should be allowed, even if it slows down the approval process? Is it the publics responsibility to remain vigilant or is governmental review of HCPs sufcient? Questions 1. What are the concerns of developers versus environmentalists with regard to natural areas? 2. Is it short-sighted to stress the importance of jobs over the rights of wildlife? Why or why not? 3. In what ethical ways can each side make their concerns known to the general public? Back Forward Main Menu TOC Study Guide TOC Textbook Website Student OLC MHHE Website Go To Student OLC 5. Why doesnt science answer ethical and moral questions? 6 6. Name eight characteristics of life, and discuss each one. 7-10 7. Food provides which two necessities for living things? 8 8. Give an example of homeostasis. Tell how the digestive sys- tem contributes to homeostasis in humans. 8 9. Why is the phrase lifelong developmental change appro- priate when speaking of humans? 9 10. How is an osprey adapted to its way of life? 9 11. How does evolution explain both the diversity and the unity of life? 9 12. What are the levels of biological organization beyond the organism? Dene each level. 10 13. Name the categories of classication, from genus to kingdom. Which category contains more types of organisms having general characteristics in common? 13 14. Explain the scientic name of an organism. 13 Testing Yourself Choose the best answer for each question. 1. Science always studies an event that a. has previously been published. b. lends itself to experimentation. c. is observable. d. ts in with an already existing theory. e. Both b and c are correct. 2. After formulating a hypothesis, a scientist a. proves the hypothesis true or false. b. tests the hypothesis. c. decides how to have a control. d. makes sure of his variables. e. formulates a theory. 3. Ascientist cannot a. make value judgments like everyone else. b. prove a hypothesis true. c. contribute to a long-standing theory. d. make use of pre-existing mathematical data. e. be as objective as possible. For questions 47, match the statements in the key with the sentences below. Key: a. Living things are organized. b. Living things are homeostatic. c. Living things respond to stimuli. d. Living things reproduce. e. Living things are adapted. 4. Genes made up of DNAare passed from parent to child. 5. Cells are made of molecules, tissues are made of cells, and organisms are made of tissues. 6. Aherd of zebra will scatter when a lion approaches. 7. The long, sharp claws of an osprey can hold on to a sh. 8. An example of chemical cycling occurs when a. plants absorb solar energy and make their own food. b. energy ows through an ecosystem and becomes heat. c. osprey nest on the top of telephone poles. d. death and decay makes inorganic nutrients available to plants. e. we eat food and use the nutrients to grow/repair tissues. 9. Which of these is mismatched? a. Kingdom Moneramosses, ferns, pine trees b. Kingdom Protistaprotozoans, algae, water molds c. Kingdom Fungimolds and mushrooms d. Kingdom Plantaewoody and nonwoody owering plants e. Kingdom Animaliash, reptiles, birds, humans. 10. An investigator spills dye on a culture plate and then notices that the bacteria live despite exposure to sunlight. He hypothesizes that the dye protects bacteria against death by ultraviolet (UV) light. To test this hypothesis, he decides to expose two hundred culture plates to UV light. One hundred plates contain bacteria and dye; the other hundred plates contain only bacteria. Result: after exposure to UV light, the bacteria on both plates die. Fill in the right-hand portion of this diagram. Chapter 1 The Study of Life 15 1-15 conclusion hypothesis experimentation and/or observations Scientific Method observations d. b. c. Example a. Thinking Scientically 1. Considering controlled experiments (page 4): a. Avariable is an element that changes. Why is sweetener S called the experimental variable in the experiment? b. With reference to Figure 1A, explain this denition: A control group goes through all the steps of an experiment except the one being tested. c. Why is bladder cancer the dependent variable in the described experiment? d. Does the experiment have elements that are constant and not variable? What are they? e. What is the value of including a control group in an exper- iment? 2. Considering the scientic process (pages 26): a. Scientic hypotheses must be falsiable. Why is the hypothesis Every human being has a guardian angel not falsiable? b. Why is the hypothesis Biotin is required for good health falsiable? c. In what way are religious beliefs different from scientic beliefs? Back Forward Main Menu TOC Study Guide TOC Textbook Website Student OLC MHHE Website 16 Introduction 1-16 Match the terms to these denitions: a. Concept consistent with conclusions based on a large number of experiments and observations. b. Statement that is capable of explaining present observations and will be tested by further experimentation and observations. c. Capacity to do work and bring about change; occurs in a variety of forms. d. Suitability of an organism for its environment enabling it to survive and produce offspring. e. Maintenance of the internal environment of an organism within narrow limits. Understanding the Terms adaptation 9 behavior 8 biodiversity 12 biology 2 biosphere 10 cell 7 community 7 conclusion 5 control group 3 data 5 development 9 ecosystem 7 energy 8 evolution 9 experiment 3 falsify 5 gene 9 homeostasis 8 hypothesis 2 law 5 metabolism 9 organ 7 organism 7 organism system 7 phenomenon 2 population 7 principle 5 reproduce 9 science 2 scientic method 6 scientic theory 5 species 9 taxonomy 13 technology 6 tissue 7 variable 3 Further Readings for Chapter 1 Balick, M. J., and Cox, P. A. 1996. Plants, people, and culture: The science of ethnobotany. New York: Scientic American Library. This well-illustrated book discusses the medicinal and cultural uses of plants, and the importance of rain forest conservation. Barnard, C., et al. 1993. Asking questions in biology. Essex: Longman Scientic & Technical. First-year life science students are introduced to the skills of scientic observation. Carey, S. S. 1997. Abeginners guide to scientic method. 2d ed. Belmont, Calif.: Wadsworth Publishing. The basics of the scientic method are explained. Cox, G. W. 1997. Conservation biology. Dubuque, Iowa: Wm. C. Brown Publishers. This text examines the eld of conservation, surveys basic principles of ecology and considers how biodiversity can be preserved. Dobson, A. P. 1996. Conservation and biodiversity. New York: Scientic American Library. Discusses the value of biodiversity, and describes attempts to manage endangered species. Drewes, F. 1997. How to study science. 2d ed. Dubuque, Iowa: Wm. C. Brown Publishers. Supplements any introductory science text; shows students how to study and take notes and how to interpret text gures. Frenay, A. C. F., and Mahoney, R. M. 1997. Understanding medical terminology. 10th ed. Dubuque, Iowa: Wm. C. Brown Publishers. Astructural approach to the study of medical terminology. Johnson, G. B. 1996. How scientists think. Dubuque, Iowa: Wm. C. Brown Publishers. Presents the rationale behind 21 important experiments in genetics and molecular biology that became the foundation for todays research. Kellert, S. R. 1996. The value of life: Biological diversity and human society. Washington D.C.: Island Press/Shearwater Books. The importance of biological diversity to the well-being of humanity is explored. Marchuk, W. N. 1992. Alife science lexicon. Dubuque, Iowa: Wm. C. Brown Publishers. Helps students master life sciences terminology. Margulis, L., et al. 1998. Five kingdoms: An illustrated guide to the phyla of life on earth. New York: W. H. Freeman & Co. Introduces the kingdoms of organisms. Minkoff, E. C., and Baker, P. J. 1996. Biology today: An issues approach. New York: The McGraw-Hill Companies, Inc. This introductory text emphasizes understanding of selected biological issues, and discusses each issues social context. Nemecek, S. August 1997. Frankly, my dear, I dont want a dam. Scientic American 277(2):20. The article discusses how dams affect biodiversity. Primak, R. B. 1995. Aprimer of conservation biology. Sunderland, Mass.: Sinauer Associates. Addresses the loss of biological diversity throughout the world, and suggests remedies. Schmidt, M. J. January 1996. Working elephants. Scientic American 274(1):82. In Asia, teams of elephants serve as an alternative to destructive logging equipment. Serani, A. 1993. The epic history of biology. New York: Plenum Press. This is a history of biology beginning with ancient Egyptian medicine. Using Technology Your study of biology is supported by these available technologies: Essential Study Partner CD-ROM Evolution & Diversity Classication Visit the Mader web site for related ESP activities. Exploring the Internet The Mader Home Page provides resources and tools as you study this chapter. http://www.mhhe.com/biosci/genbio/mader Back Forward Main Menu TOC Study Guide TOC Textbook Website Student OLC MHHE Website