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189
Introduction
Throughout most of the Sonoran Desert the dominant
woody species are members of the Leguminosae. Exceptions to
this lie in the northwestern part of the desert where relatively
cool winters and low, cool season rainfall apparently conspire
190
63
7(6).
Olneya)
8
Leaves odd -pinnate (terminal leaflet
7.
8(7).
8.
glabrous
10(8).
10.
11(10).
the key.
11.
Key
2(1).
2.
2
6
12(10).
Psorothamnus spinosus
4.
Desmodium
psilophyllum Schlecht.
13.
14(12).
14.
15.
5.
6(1).
6.
15(14).
5(4).
13
12
Shrubs
Leaflets 2 -5 pairs, large, to 7 cm wide
and 15 cm long, persistently tomentulose beneath, margins revolute; twigs
Cassia
pubescent, straight
emarginata L.
Leaflets 8 -20 pairs, 1 -2 cm long,
glabrous, margins not revolute; twigs
glabrate, zig -zag
Tamarindus
indicus L.
13(12).
Haematoxylon
brasiletto Karst.
gland(s)
12.
3(2).
9
10
9.
Leaves simple
Leaves once or twice pinnate
17
9(8).
rachis (Papilionoideae)
Plants armed
Plants unarmed
24(23).
Errazurizia
megacarpa
(S. Wats.) I. M. Johnston
26
Branches spinescent
caducous
4 -9
17(7).
17.
191
25(23).
25.
fremontii)
26(25).
veined, to 9 cm long
Eriosema grandiflorum (Schl. &
Brandegee
Cham.) Seem.
18.
26.
19(18).
20(19).
27.
wide
Psorothamnus arborescens
var. simplifolius
trifoliata Brandegee
Plants suffrutescent, to 1.5 m high;
28(25).
Eysenhardtia orthocarpa
mainland Mexico
Desmodium cinerascens A. Gray
21.
28.
22.
30
29.
30(29).
30.
caducous
1/
23(22).
2/
29
22
'/
29(28).
(Parish) Barneby
Leaflets mostly 21 -51
1/
20
Spreading shrub, 2 -4 m high; leaf-
1/
27/26).
19.
28
31(29).
1/
192
Psorothamnus emoryi
(A. Gray) Rydberg
Terminal leaflets not distinctly longer
32
than the lateral ones
31.
32(31)
32.
`/
2/
40(22).
40.
34
33.
41(40).
41.
42(41).
42.
(Torr.) Barneby
Leaflets 1 -5 mm long; petioles shorter
Dalea pulchra
Gentry
36(33).
1/
Rydb.
43.
1/
(Rose) Bullock
Foliage with dense to sparse pubes-
base
44(43).
37
S. Wats.
44.
Dalea scandens
(Miller) R. T. Clausen
var. occidentalis (Rydb.) Barneby
38(37).
45(42).
38.
brous to pilosulous
Dalea versicolor Zucc.
var. sessilis (A. Gray) Barneby
Petioles longer; young branches
Rose
Bark grey, smooth; leaflets oblong elliptic, 1.0 -1.5 cm wide and 2.4 cm
45.
1/
silky -pilosulous
39
Stipules 0.8 -1 5 mm long, puberulent,
triangular -subulate; leaflets
(1- )2 -3( -4) pairs + 1; intra- petiolular
1/
37.
44
1/
43
45
Psorothamnus fremontii
35.
Leaves estipulate
43(42).
35(34).
34.
1/
33(32).
Willardia mexicana
(S. Wats.) Rose
46(41).
46.
1/
47(46).
subulate
49
Leaflets oblong to oblanceolate, 1 -2
cm long, 2 times longer than broad,
3/
53.
54(53).
48(47).
Rydb.
54.
55(54).
56(40).
56.
57(56).
55.
48.
Indigofera suffruticosa
Mill.
Branches terete
54
Leaflets 5 in number
Indigofera laevis
3/
47.
193
R. Br.
57.
58(57).
Rose
49(46).
49.
Leaflets glabrous
50
Leaflets pubescent, at least on lower
surface (occasionally glabrate in age)
59.
60(59).
S. Wats.
50.
52(51).
61
Branches angled
strongly 3 -ribbed
Tephrosia leiocarpa
A. Gray
52.
61(60).
Pubescence of malpighiaceous
51.
Rose
Leaflets and branchlets not glandular viscid
60
Leaflets silvery sericeous
Brandegee
60.
51(49).
Diphysa racemosa
Aeschynomene nivea
59
59(58).
51
50(49).
62.
194
63(6).
Mimosa margaritae
Rose
73.
64
only
63.
may be spinescent
64(63).
103
74(73).
74.
29 30' N
.
Prosopidastrum mexicanum
65(64).
65.
66(65).
65
66
less)
81
76 (75).
76.
67(66).
67.
68
69(68).
prickles
70
Pinnae (1 -)4 -18 pairs, leaflets
linear
Acacia kelloggiana
Carter and Rudd
Pinnae 1 -4 pairs, leaflets obovate
or oblong
69
77.
4/
71(70).
spines
Pithecellobium mexicanum Rose
Gland near middle of
petiole
71
Gland between lower pair of
pinnae
72
Petioles 4 -6 mm long; leaflets 5 -15
pairs, 0.6 -1 mm broad, 3 -5 mm long;
Sonora
Acacia occidentalis
Benth.
78.
Rose
Rose
71.
79(76).
72.
79.
80(79).
73(66).
Pinnae
6 -14 mm long
Acacia wrightii
Benth.
mm long
78
based spines
Pithecellobium sonorae S. Wats.
Leaflets 3- 10( -12) pairs, midvein
69.
68
68(67).
(L.) Swartz
76
77
79
77(76).
67
66.
75.
74
75(74).
(Dressl.) Burk.
64.
above
Mimosa brandegeei
Robinson
5/
80.
81(65).
81.
82(81).
82.
83(82).
...
Mimosa purpurascens
Robinson
Petioles without glands
82
Petioles with glands
88
Bark of large branches green or
greenish
83
Bark otherwise
87
Leaves 3- 30( -60) cm long; leaflets
long
89.
84(83).
90(88).
91
91.
S. Wats.
1( -3)
92(91).
long
85.
Cercidium praecox
(Ruiz & Pay.) Harms
92.
wise
6/
94
93.
86.
94.
8/
95(94).
88(81).
88.
10/
u0/
87.
long as broad
Prosopis glandulosa Torr.
var. torreyana
(L. Benson) M. C. Johnston
Leaflets less than 5 times as long
as broad
95
Foliage puberulous or glabrous; pinnae 1 pair on short shoots and 1.0 -2.5
cm long; pinnae 1 or 2 pairs on long
shoots and 2.5 -4 cm long
Prosopis articulata
Foliage markedly short -pubescent to
S. Wats.
95.
(Rose) Carter 8/
87(82).
93
9/
93(92).
86(84).
91(90).
85
(Roxb.) Benth.
Leaflets 2(3) pairs, 1 -2.5 cm long
Acacia californica
Brandegee
Pinnae 30 -60 pairs; leaflets (30 -)
90
Pithecellobium dulce
83.
195
96.
shoots
96
Leaflets with 3 -4 prominent veins
arising from base; petiolar gland
near middle of petiole
Pithecellobium undulatum
(Britt. & Rose) Gentry
11/
89
Prosopis velutina
Wooton
10/
196
105.
98
97(93).
97.
106(105).
110
Leaflets smaller
Leaves with glands on rachilla
107
99
98(97).
107(106).
12/
107.
Acacia constricta
99(97).
13/
Pithecellobium confine
Standley
99.
100.
S. Wats.
109(106).
109.
Brandegee
101(100).
101.
Leucaena microcarpa
Rose
110(105).
110.
Prosopis palmen
Lysiloma candida
Brandegee
10/
104.
125
Caesalpinia)
Petioles phyllode -like, 1.5 -4 mm
wide, 6 -30 cm long
Acacia willardiana
Rose
Petioles not as in foregoing
105
105(104).
111
112
S. Wats.
Petioles 5 -9 mm long; short shoots
103(63).
103.
111(110).
between 24 N and 27 N
102.
Benth.
and
A. pacensis
113(112).
113.
D. covillei
(Britt. & Rose) Wiggins ex Turner
Trees
113
112(110).
117
112.
Shrubs
114
Petioles 2.5 -5 cm long
115
Petioles 0.5 -2.5 cm long
114(113).
114.
14/
Acacia coulteri
Benth.
pairs of pinnae
Desmanthus subulatus
(Britt. & Rose) Wiggins ex Turner
115(113).
Pithecellobium tortum
Mart. u/
115.
124.
197
nae
Desmanthus virgatus
116.
Willd.
125(103).
125.
Rose
Leaves 6- 10( -12) cm long; pinnae
126
127
14/
126(125).
Torr.
126.
Mart.
Petiolar glands between or just below
lowermost pair of pinnae, or, if near
mid -petiole, between pair of bracts
117.
11/
127(125).
118
118(117).
118.
119(118).
119.
Wall.
127.
Lysiloma watsonii
121(120).
121.
Rose
Leaves 6- 10( -12) cm long; pinnae
121
2.5 -4.5 cm long
Pinnae 4 -10 pairs; leaflets 20 -30 pairs
Lysiloma microphylla
Benth.
122.
123.
130(129).
Caesalpinia placida
Brandegee
Rose
Lysiloma
129.
Desmanthus fruticosus
....
123(122).
10./
pairs
122(118).
128
129
132
129(128).
lent
120
120(119).
Leaves 10 -20 cm long; pinnae
120.
14/
117(112).
124
130.
14/
131(130).
mm long
131
Pinnae 1 -2( -3) cm long; leaflets
5- 12( -15) mm long
Caesalpinia palmeri
S. Wats.
and
C. arenosa
Wiggins
198
131.
132.
Cercidium microphyllum
(Torr.) Rose & Johnston
Leaves petiolate, usually longer;
pinnae 1 or more pairs; leaflets
133
mostly longer
133(132).
133.
134
maturity
Distal pinnae shorter when mature
Benth.
140(137).
Acacia crinita
Brandegee
140.
141(140).
Brandegee
142
142(141).
glabrous
Caesalpinia
platyloba S. Wats.
143(142).
135(133).
135.
136
136( 135).
136.
137( 136).
137.
138(137).
Brandegee
144(135).
144.
(L.) Swartz
137
145)144).
138
shorter than 1 cm
Leaves stipulate or estipulate; petioles longer than 1 cm if shorter,
140
leaves estipulate)
145.
(Boj.) Raf.
146.
139
139.
Calliandrd eriophylla
Benth.
Stipules 2 -3( -7) mm long; petioles
4 -10 mm long; pinnae 1 -2) -4) pairs,
1.5 -4 mm long; found only in Baja
Torr. ex S. Wats.
145
Trees
146
Shrubs
Pubescent, spheroidal projections be-
Rose
Leaflets 5 -15 pairs, 1.5 -3 times as
Acacia crinita
143.
Not as above
hispid
138.
densely so beneath
Conzattia sericea Standley
134.
141
Twigs not as above
Leaflets 2 pairs, dimorphic (adaxial
leaflet of lower pair usually reduced
to 2 -7 mm long, other leaflet 1 -3 cm
long)
Cal liandra rupestris
141.
135
134(133).
A. mcmurphyi
Wiggins
and
A. angustissima
(Mill.) Kuntze
15/
Footnotes
1.
2.
3.
Blade
Bristles
Stiff hairs.
Caducous
Canescent
Cauline
Ciliate
5.
199
hairs.
Cinereous
(1980) lists this species for east -central Baja California. We know of no
Cla va to
Compound
Confluent
Cordate
Coriaceous
Leathery in texture.
Crateriform
Crenulate
Deciduous
Decurrent
Distal
Glossary
terminal.
Divaricate
Widely spreading.
Eccentric
Off -center.
Elliptic
Emarginate
Epidermis
Estipulare
Lacking stipules.
Even -pinnate
Fascicle
Filiform
Thread -like.
Fissured
Cracked.
Acicular
Needle- shaped.
Glabrate
Acute
Glabrous
Without hairs.
sharp point.
Glandular
Abaxial
Adaxial
axis; ventral.
Alternate
Glaucous
Herbage
Annular
Hirsute
Apiculate
Hispid
Appressed
Imbricate
Internode
leaf.
Armature
Armed
Axillary
Isodiametric
lunceous
Oval
Lana te
Lanceolate
Leaflet
Lenticel
Linear
Ovate
Papillose
Pellucid
Translucent, clear.
Persistent
Petiole
Petiolule
Malpighiaceous
Phyllode
Mucronate
With a short, abrupt (but not sharp), hardened point or mucro; compare pungent,
Phylloidal
Leaf -like.
apiculate.
Pilose
Node
Oblanceolate
Pilosulous
narrow base.
Pinna(e)
Pinnate
Oblong
Obovate
Obtuse
Odd -pinnate
Pithy
Soft, spongy.
Prickle
Once pinnate
Primary leaves
Opposite
Puberulent
Puberulous
Orbicular
Circular in outline.
pubescent.
INTRA-PETIOLULAR
GLAND
EVEN
PINNATE
LEAF
ODD
PINNATE
LEAF
PETIOLULE
STIPULE
ONCE
PINNATE
LEAF
POST- PETIOLULAR
GLANDS
Figure 3. Segment of Olneya tesota stem showing once pinnate leaves with odd- pinnate and even -pinnate arrangement
of leaflets.
The leaves are once -pinnate with even -pinnate and odd pinnate arrangement of leaflets. The leaflets and stem are
glandular punctate.
Pubescent
Punctate
Pungent
(Fig. 4).
5,6).
mucronate.
Pustules
Small blisters.
Pustulate
Rachilla
Spinescent
Somewhat spiny, or becoming spiny; terminated by a sharp tip or spine (Fig. 2).
Spinose
Having spines.
Stipel
Stipellate
Having stipels.
Stipitate
Rachis
Spine
sessile.
Recurved
Remote
Reticulate
Striate
Retuse
Strigose
Revolute
Strigulose
Rhombic
Sub -nodal
Sericeous
Subulate
Sessile
Setaceous
Suffrutescent
Setiform
Sulcate
Grooved or furrowed.
Short shoot
Terete
Tree
having several stems from the ground; without a single main trunk.
Tomen tose
Tomentulose
Shrub
Simple
Stipule
hairs.
Tomentum
Tuberculate
Twice- pinnate
Describes a compound leaf with both primary and secondary divisions pinnate (Fig.
1,2,5,6).
Velutinous
Venation
Verruculose
Villous
Viscid
Sticky.
1974b. The genus Cercidium (Leguminosae: Caesalpinioideae) in the Sonoran Desert of Mexico and the United States.
Proceedings of the California Academy of Science 40(21:17-57.
Felger, Richard S., and Lowe, Charles H. 1970. New combinations for
Literature Cited
Barneby, Rupert C. 1977. Daleae imagines, an illustrated revision of
Errazurizia Philippi, Psorothamnus Rydberg, Marina Liebmann, and
Dalea Lucanus emend. Bameby, including all species of Leguminosae
and Carter, Annetta M. 1983. Acacia pacensis (Leguminosae: Mimosoideae), a new species from Baja California del Sur,
Mexico. Madrollo 30(3): 176 -180.
Shreve, Forrest. 1964. Vegetation of the Sonoran Desert. Vol. 1, Part 1. In