Professional Documents
Culture Documents
R & D department
TROPICA SEEDS PVT LTD | No 54,
South End Road, 1st Floor, Nama
Aurore Building, Basavangudi,
Bangalore 560004 INDIA
Page
Disclaimer
This publication may be of assistance to you but Tropica seeds and its employees
do not guarantee that the publication is without flaw of any kind or is wholly appropriate
for your particular purposes and therefore disclaims all liability for any error, loss or other
Consequence which may arise from you relying on any information in this publication.
Page
II
Contents
Page no.
1-2
3
Bacterial diseases
4
Xanthomonas campestris pv. vesicatoria
5
Ralstonia solanacearum
Clavibacter michiganensis subsp. michiganensis 6
Fungal diseases
Colletotrichum gloeosporioides
Cercospora capsici
Choanephora cucurbitarum
Fusarium oxysporum f.sp. capsici
Fusarium solani
Stemphylium solani
Botrytis cinerea
Phytophthora capsici
Leveillula taurica
Sclerotium rolfsii
Verticillium spp
Sclerotinia sclerotiorum
7
8
9
10
11
12
14
15
16
17
Abiotic diseases
Blossom-end rot
Sunscald
28
29
18
ReferencesMeloidogyne incognita
20
21
22
23
24
25
26
27
13
Nematodes, parasitic
Root knot
Alfalfa mosaic
Alfalfa mosaic virus (AMV)
Beet curly top
Beet curly top virus (BCTV)
Chili leaf curl
Tobacco leaf curl virus (TLCV)
Cucumber mosaic
Cucumber mosaic virus (CMV)
Potato virus Y
Potato virus Y (PVY)
Tobacco etch
Tobacco etch virus (TEV)
Tobacco mosaic and Tomato mosaic Tobacco mosaic virus (TMV)
Tomato spotted wilt
Tomato spotted wilt virus (TSWV)
30
19
III
Anthracnose
Cercospora (frogeye) leaf spot
Choanephora blight (wet rot)
Fusarium wilt
Fusarium stem and root rot
Gray leaf spot
Gray mold
Phytophthora blight
Powdery mildew
Southern blight
Verticillium wilt
White mold
Viral diseases
Page
Bacterial spot
Bacterial wilt
Bacterial canker
This article presents the various steps/activities which are associated with
accurate plant disease diagnosis. The process may vary with different diseases and
conditions but the overall process is relatively consistent. The steps all require
careful observations and questions. The steps include:
Know what Normal is
Proper plant identification. Identification of affected plants is one of the first
steps in diagnosing a plant disease. Both scientific and common names of the plant
should be noted.
Recognize healthy plant appearance. It is important to know the normal
appearance of the plant species you are investigating. Each plant species has
special growth habits, colours and growth rates. If you do not know what to expect
of the plant you cannot recognize when something is wrong.
Page
The diagnostician must have very good observation skills, and he or she also
needs to be a good detective. It is important to keep an open mind until all of
the facts related to the problem can be collected. The possibility of multiple
causal factors must also be considered. Control measures depend on proper
identification of diseases and of the causal agents. Therefore, diagnosis is one
of the most important aspects of a plant pathologist's training. Without
proper identification of the disease and the disease-causing agent, disease
control measures can be a waste of time and money and can lead to further
plant losses. Proper disease diagnosis is therefore vital.
Check for host specificity. Is the problem occurring in only one plant species or
are different plant species affected? If different plant species are affected, this
suggests the possibility of a non-infectious problem which could be related to
cultural or environmental problems. However, Phytophthora and Pythium root
rots can cause problems on many different plant species; therefore, the fact that
more than one plant species is affected does not completely eliminate infectious
agents. If there is more than one species of plant involved, are these plants closely
related and can they be infected by a common pathogen?
Laboratory Tests
Sometimes neither symptoms nor signs provide enough specific or characteristic
information to decide the cause of an infectious plant disease. In such cases, it may
be necessary to bring a sample back to the laboratory for further tests to isolate
and identify the causal agent.
Incubation of plant material. One of the first steps when getting back to the
laboratory may be to place a sample of the diseased tissue under conditions that
will allow an infectious agent to grow and possibly induce sporulation. This can be
accomplished by placing a leaf in a moist chamber. A moist chamber can be a
Page
Identify Plant Part Affected - Are symptoms associated with specific plant
parts?
It is important to note if the symptoms observed are associated with specific plant
parts. For example, is a wilt observed correlated with a disruption of the vascular
system which may be indicated by browning of the vascular system or are the roots
of the plants abnormal including rots, decreased feeder roots, etc.; are necrotic
lesions observed strictly on younger leaves? The symptoms of some diseases are
most commonly seen on specific plant parts and this observation can be important
in diagnosis.
sterile petri dish containing a wet filter paper in the bottom of the dish and a
triangle of glass tubing on which the sample is placed so that the sample is not
directly on the wet filter paper but is exposed to humid conditions. This type of
moist chamber will work for small and relatively flat specimens such as leaves.
Plastic bags or boxes may be necessary for larger specimens. Saprophytes that are
present on the specimen can also be encouraged to grow in a moist chamber and a
brief surface swab with 70% isopropanol or 0.1-1% sodium hypochlorite may be
useful in reducing these saprophytes. Moist chambers are generally incubated at
room temperature.
EXCLUSION
This principle is defined as any measure that prevents the introduction of a
disease-causing agent (pathogen) into a region, farm, or planting. The basic
strategy assumes that most pathogens can travel only short distances without the
aid of some other agent such as humans or other vector, and that natural barriers
like oceans, deserts, and mountains create obstacles to their natural spread.
RESISTANCE
Use of disease-resistant plants is the ideal method to manage plant diseases, if
plants of satisfactory quality and adapted to the growing region with adequate
levels of durable resistance are available. The use of disease-resistant plants
eliminates the need for additional efforts to reduce disease losses unless other
diseases are additionally present.
ERADICATION
This principle aims at eliminating a pathogen after it is introduced into an area but
before it has become well established or widely spread. It can be applied to
individual plants, seed lots, fields or regions but generally is not effective over large
geographic areas.
The goal of plant disease management is to reduce the economic and aesthetic
damage caused by plant diseases. Specific management programs for specific
diseases are not intended since these will often vary depending on circumstances
of the crop, its location, disease severity, regulations and other factors. Plant
disease management practices rely on anticipating occurrence of disease and
attacking vulnerable points in the disease cycle (i.e., weak links in the infection
chain). Therefore, correct diagnosis of a disease is necessary to identify the
pathogen, which is the real target of any disease management program.
PROTECTION
This principle depends on establishing a barrier between the pathogen and the
host plant or the susceptible part of the host plant. It is usually thought of as a
chemical barrier, e.g., a fungicide, bactericide or nematicide, but it can also be a
physical, spatial, or temporal barrier. The specific strategies employed assume that
Page
The many strategies, tactics and techniques used in disease management can be
grouped under one or more very broad principles of action. Included four general
disease control principles, exclusion,eradication, protection&Immunization(the
latter principle is more appropriately called resistance since plants do not have an
immune system)
pathogens are present and that infection will occur without the intervention of
protective measures.
Many cultural practices can be modified to manage the occurrence, intensity or
severity of plant diseases. These include selection of suitable growing sites for the
crop, adequate tillage to bury pathogen-infested plant residues, rotation to non
susceptible crops, selecting pathogen-free planting stocks, orientation of plantings
to improve exposure to sun and air currents, pruning and thinning to eliminate
sources of infection and improve aeration in and around susceptible plants, water
management on both plants and in soil, adequate nutrition, proper cultivation to
improve root growth and avoid plant injury, and sanitation procedures to eliminate
sources of inoculum.
Biological control involves the use of one living organism to control another, and
this management technology has received much attention in recent times. However,
the number of biological agents registered for use is relatively small, success has
been limited, and application has been largely restricted to intensively managed,
high value crops such as greenhouse plants.
Chemical control
Chemical name
Xanthomonas is seed-borne on
the seed surface and within
the seed. Infected seed and
transplants moved over long
distances can be the initial
source of inoculum for
epidemics. The bacterium also
survives in crop debris,
volunteer plants and in
solanaceous
weeds.
High
relative humidity and heavy
dew formation on leaves,
together with warm weather,
favour
infection
and
development of Bacterial Spot.
The bacterium is readily
water-splashed from infected
Transplants or debris to
healthy plants. Fruit are
infected
through
growth
cracks,
abrasions,
insect
punctures and other wounds.
Secondary fruit rots often
develop around Bacterial Spot
lesions during damp weather.
Dosage(g
PHI
or ml/litre) (days)
Remarks
Copper hydroxide
(kocide)
2g/litre
Copper sulphate
(cuproxat)
2g/litre
Streptomycin
sulphate+ tetracyclin
hydrochloride
(agrimycin)
Mancozeb
(Dithane M-45)
6g/10 litre
15
3g/litre
10
Should be used on
younger stage of the crop
growth, do not use
during fruiting stage
Used as a protectant
Cuprous oxide
(Nordox)
2g/litre
Note:
1. Copper fungicides should not be used at flowering stage in
solanaceous crops.
2. Before using combination sprays, please check the label of the
product
Page
Symptoms
Chemical control
Chemical name
Dosage(g or
ml/litre)
PHI
Remarks
Streptomycin sulphate+
tetracyclin hydrochloride
(Agrimycin)
6g/10 litre
15
Should be used on
younger stage of the
crop growth, do not use
during fruiting stage
Copper oxychloride
(Blitox)
3g/litre
Initial symptoms
of wilt
occur in younger leaves and
slight yellowing of older
leaves. The wilted leaves
maintain their green colour
and do not fall off as disease
progresses. Under conditions
favourable
to
the
disease Complete wilt occurs.
Wilting
and
death
is
accompanied by dark brown
internal discoloration of the
vascular elements.
White, milky strand of
bacterial
ooze
flows out when a freshly cut
section of infected stem base
is placed in water.
Page
Symptoms
Chemical control
Chemical name
Streptomycin sulphate+
tetracycline hydrochloride
(agrimycin)
Copper oxychloride
(Blitox)
Dosage(g
or
ml/litre)
6g/10
litre
PHI
Remarks
15
3g/litre
Should be used on
younger stage of the crop
growth, do not use
during fruiting stage
Used as soil drenching at
the root zone of the plant
Note:
1.
Copper fungicides should not be used at flowering stage in
solanaceous crops.
2.
Before using combination sprays, please check the label of the product
Page
Symptoms
Anthracnose of Pepper
Causal organism: Colletotrichum capsici
Chemical control
Chemical name
Dosage(g
PHI
or ml/litre) (days)
Plant only seed from disease- Mancozeb+carbendazim 2+1g/litre 3
freeplants or seed treated to reduce any (Dithane M-45+
fungal populations. Seed can be Bavistin)
disinfested with a 30-minute soak at
52C. Use only transplants free Pyraclostrobin (cabrio) 1g/litre
0
of disease symptoms.
Crop rotation Fields should be planted
with crops other than tomatoes,
eggplants, or other solanaceous crops
Chlorothalonil (kavach) 2g/litre
3
or strawberries, which are also hosts.
Apply overhead irrigation during the
early part of the day so that plants can
dry before sundown.
Azoxystrobin (amistar) 0.5ml/litre 0
In areas where market constraints and
other diseases do not limit the choice of
cultivar, cultivars demonstrating a
moderate level of resistance (e.g.,
Colossal, Brigadier, and Paladin) should
Trifloxystrobin+
1g/litre
3
be chosen.
Tebuconazole(nativo)
Remarks
Used as a protectant as
well as curative action
Note:
For mixing chemical always see the label of the product.
Page
Symptoms
Chemical control
Chemical name
Dosage(g
PHI
or ml/litre)
Remarks
Chlorothalonil
(kavach)
2g/litre
Can be used as a
protective application;
5-7 days interval
Mancozeb
(Dithane M-45)
2g/litre
Used as a protective
application; 5-7 days
interval
Copper hydroxide
(kocide)
2g/litre
Used as protective
application
Trifloxystrobin+
Tebuconazole
(nativo)
1g/litre
Used as curative
applicant
Note:
For mixing chemical always see the label of the product.
Page
Symptoms
Chemical control
Chemical name
Dosage(g
or ml/litre)
PHI
Remarks
Chlorothalonil(kavach)
2g/litre
Can be used as a
protective application;
5-7 days interval
Mancozeb
(Dithane M-45)
2g/litre
Used as a protective
application; 5-7 days
interval
Propineb
(Antracol)
2g/litre
Trifloxystrobin+
Tebuconazole
(nativo)
1g/litre
Used as curative
applicant
Myclobutanil
(Systhane)
1g/litre
14
Note:
For mixing chemical always see the label of the product.
Page
Symptoms
Chemical control
Chemical name
Dosage(g
PHI
or ml/litre)
Remarks
3g/litre
1g/litre
10
Page
Symptoms
h.
i.
j.
k.
l.
Discard slabs, bags, cubes or other media that had infected plants growing in
them previously.
Do not replant into the same material unless it has been steam-sterilized.
Remove and discard strings that may harbour spores from affected plants.
If the crop was grown in soil, disinfect the beds.
Discard soilless growing media far away from the greenhouse or bury it.
Avoid handling diseased plants and fruit.
Remove them from the greenhouse carefully, taking care not to allow
contact of affected portions of plants with adjacent plants, and place
them in a plastic bag.
Discard the diseased material at a location away from the
greenhouses to ensure that this fungus inoculum or subsequent
overwintering spore inoculum is not carried back into the
greenhouses by workers, wind, on tires, and by insects such as shore
flies and fungus gnats. Additionally, remove about 1-2 plants on
either side of the plant(s) exhibiting symptoms and place in garbage
bags.
If the material is disposed of in a cull pile then ensure that the cull
pile is located away from the greenhouse as far as possible.
Cover the cull pile to prevent insects such as shore flies and fungus
gnats from carrying the fungus spores back into the greenhouses.
Alternatively, infected plant debris may be burnt or taken to a
landfill.
Do not leave it in an open field or incorporate it into the soil in fields
where other susceptible crops may be grown
11
Page
Symptoms
Chemical control
Chemical name
Dosage(g
PHI
or ml/litre)
Remarks
Chlorothalonil
(kavach)
2g/litre
Mancozeb
(Dithane M-45)
2g/litre
Propineb
(Antracol)
2g/litre
Can be used as a
protective application;
5-7 days interval
Used as a protective
application; 5-7 days
interval
5-7 days interval
Difenoconozole
(Score)
0.5ml/litre
10
Used as curative
fungicide
Copper hydroxide
(kocide)
2g/litre
Used as a protective
application
Tebuconazole
(Folicur)
1ml/litre
Used as a curative
application
Note
For tank mixing of different chemicals see the label.
12
Page
Symptoms
Chemical control
Chemical name
Dosage(g
or
ml/litre)
PHI
Remarks
Iprodione+carbendazim
(quintal)
1g/litre
Chlorothalonil
(kavach)
2g/litre
Can be used as a
protective application;
5-7 days interval
Pyraclostrobin
(Cabrio)
1g/litre
Difenoconozole
(Score)
0.5ml/litre
10
Used as curative
fungicide
Copper hydroxide
(kocide)
2g/litre
Used as a protective
application
Pyrimethanil
(scala)
0.5ml/litre
Repeat applications at
7-14 day intervals
Note
For tank mixing of different chemicals see the label.
13
Page
Symptoms
Phytophthora blight
of Peppers
Causal organism: Phytophthora capsici
Chemical control
Chemical name
Dosage(g
PHI
or ml/litre)
Remarks
Mancozeb
(Dithane M-45)
2g/litre
Used as a protective
application; 5-7 days
interval
Chlorothalonil
(kavach)
2g/litre
Can be used as a
protective application;
5-7 days interval
Propineb
(Antracol)
2g/litre
Dimethomorph
(Acrobat)
1g/litre
Metalaxyl+mancozeb
(Ridomil MZ gold)
2g/litre
14
Azoxystrobin
(Amistar)
0,5ml/litre
Note
For tank mixing of different chemicals see the label.
14
Page
Symptoms
Chemical control
Chemical name
Dosage(g
PHI
or ml/litre)
Remarks
Wettable sulphur
3g/litre
Dinocap
(karathane)
1ml/litre
Used as contact
fungicide, 5-7 days
application interval
Myclobutanil
(Systhane)
1g/litre
14
Fluzilazole
(nustar)
2ml/10litre
15
Used as curative
application, should not
be used at flowering
stage
For preventive
application apply once in
15 days
Carbendazim
(Bavistin)
1g/litre
Note
For tank mixing of different chemicals see the label.
15
Page
Symptoms
Chemical control
Chemical name
Dosage(g
PHI
or ml/litre)
Remarks
Carbendazim
(Bavistin)
2g/litre
Used as drenching
Copper hydroxide
(kocide)
2g/litre
Used as drenching
Fluzilazole
(nustar)
2ml/10litre
15
For preventive
application apply once in
15 days
Tebuconazole
(Folicur)
1ml/litre
For preventive
application and curative
Note
For tank mixing of different chemicals see the label.
16
Page
Symptoms
Chemical control
Chemical name
Dosage(g
PHI
or ml/litre)
Remarks
Carbendazim(bavistin)
2g/litre
Used as drenching
Copper
hydroxide(kocide)
2g/litre
Used as drenching
Note
1) For tank mixing of different chemicals see the label
2) Soil fumigation with effective materials is the only chemical control
available for reducing soilborne populations of the pathogen.
17
Page
Symptoms
Chemical control
Chemical name
Plant
surfaces
must
remain
continuously wet for 48- 72 hours for
infection to occur. Disease develops
most rapidly at 20- 25C (68- 77F),
and not at all above 30C (86F).
This fungus has a wide host range and
survives from one season to the next
as sclerotia in soil and in plant debris.
White mold is most common in
temperate regions but is also known
to occur in hot, dry areas. Dew, fog
and frequent rain generally favour
disease development. The most
important means of longdistance
spread are airborne ascospores that
erupt
from
sclerotia.
Moving
contaminated soil and fertilizing with
manure from animals fed infected
plant debris are two common ways of
short-distance spread of sclerotia or
mycelium. Irrigation water may also
spread thefungus from field to field.
Dosage(g
PHI
or ml/litre)
Remarks
Carbendazim(Bavistin)
2g/litre
Used as drenching
Copper
hydroxide(kocide)
2g/litre
Used as drenching
Flusilazole(nustar)
2ml/10litre
15
For preventive
application apply once in
15 days
Tebuconazole (Folicur)
1ml/litre
For preventive
application and curative
Note
For tank mixing of different chemicals see the label
18
Page
Symptoms
Chemical control
Chemical name
Dosage(g
or
ml/litre)
Methyl bromide
PHI
Remarks
As a
fumigant
15
Oxamyl
(vydate)
2ml/litre
14
Carbofuron
(Furadan)
4kg/hac
30
Preplant, tarped, or
mulched for 24-48
hours. Application 10=14
days before planting.
Foliar applications are not
effective for moderate and
high populations of
nematodes.
Soil application,
application into soil before
planting.
Fenamiphos
(nemacure)
30L/hac or
1L/100
litre of
water
7 days
Soil application Apply
prior to anytime from 7 days
planting before up to the time of
planting.
Note
For tank mixing of different chemicals see the label
19
Page
Symptoms
Symptoms
Control measures
Pepper varieties resistant to AMV are not available. Various control measures
are required because AMV is transmitted by seed, aphids, and mechanically.
Control measures must take into account the diseases wide host range (alfalfa,
pepper, tomato, tobacco, potato, clover, many cucurbits and beans, and several
other crops and weeds) and numerous aphid vectors.
Use virus-free pepper seed. Check transplants for any symptom development
and discard those with symptoms. Aphid control may be difficult because the
virus is transmitted very rapidly by these insects. Use fast acting insecticide
sprays since aphids may move to other nearby unsprayed plants when
disturbed.
20
Disinfect tools, stakes, and equipment before moving from diseased areas to
healthy areas. Work in diseased areas last, after working in unaffected parts of
a field. Other less effective measures include: planting barrier crops that are not
susceptible to AMV such as corn, applying sticky traps, or covering the ground
with an aphid deterrent material Another control strategy is to grow trap crops
nearby that attract aphids and then spray these plants with a contact
insecticide to destroy the aphid populations. Also, spray the pepper crop with
mineral oil to delay virus spread in the field by interfering with aphid
transmission of the virus.
Page
control measures
This virus has a wide host range affecting more than 300
species. Common hosts are tomatoes, beets, peppers, squash,
beans, cucurbits, spinach, potatoes, cabbage and alfalfa. The
beet leafhopper transmits BCTV in a persistent manner.
Warm temperatures and dense leafhopper populations are
conducive to the spread of BCTV. Viruliferous leafhoppers
migrate seasonally and can be moved long distances by wind.
Page
21
control measures
22
Page
Symptoms
Cucumber mosaic on
peppers
Causal agent: cucumber mosaic virus
control measures
Current control measures for CMV are mainly preventive due to the
wide host range of the virus and the numerous aphid vectors.
Grow seedlings in a structure or seedbed protected with netting of mesh
size of 32 or greater to prevent aphids from entering.
Disinfect tools, stakes, and equipment before moving from diseased areas
to healthy areas. Hands and tools may be washed with soap or milk.
Insecticide sprays that are not fast-acting may not be that effective
because the aphids move to other nearby unsprayed plants when
disturbed.
Other less effective measures include: planting barrier crops that are not
susceptible to CMV such as corn, applying sticky traps, or covering the
ground with an aphid deterrent material Another control strategy is to
grow trap crops nearby that attract aphids and then spray these plants
with a contact insecticide to destroy the aphid populations.
Page
Symptoms
23
Transmission: Aphids
PVY on Peppers
Causal agent: Potato virus Y
Control measures
Page
Symptoms
24
Transmission: Aphids
control measures
Use a net house or screen house with 32-mesh or finer to keep out aphids and to
grow seedlings for transplant use.
Use virus-free transplants. Minimize plant handling during the growing season to
reduce the amount of virus spread mechanically.
Remove nearby volunteer plants and solanaceous weeds from production fields,
nearby ditch banks, hedges, fence-rows or other locations. Plant earlier to avoid
high aphid populations that occur later in the season. Mulches to reduce aphid visits
to plants and thus delay virus spread. This results in reduced virus incidence and
increased yields. Monitor aphid populations early in the season and apply fastacting insecticide treatments when needed.
The mineral oil sprays will reduce the frequency of transmission of the virus by the
vector and thereby delay development of the disease in the pepper crop
Page
Symptoms
25
Transmission: Aphids
TMV on Peppers
Causal agent: Tobacco mosaic virus
control measures
TMV and ToMV have very wide host ranges and infect
over 200 plant species, including varieties of pepper,
tomato, eggplant and tobacco. TMV can infect all species
of pepper, but not tomato, tobacco or eggplant. Infected
transplants, seed and debris are common sources of
inoculum. These viruses can be found on and under the
seed coat and in the endosperm. Tobamo viruses are
very stable and extensive spread can occur through
handling, tools, trays, pots, stakes, twine and clothing, as
well as pollination, pruning and other cultural practices.
Tobamo viruses can remain viable for several years in
plant debris but lose their ability to infect as debris
decomposes.
Page
Symptoms
26
Transmission: Mechanical
control measures
The presence of thrips in pepper fields can be monitored using yellow sticky cards. If the
disease appears in a crop, infected plants should be removed and destroyed immediately
either by burning or burying them.
Maintain seedbeds away from cropped areas and from other susceptible plants.
Remove volunteer plants and weeds from the production field and surrounding areas by
maintaining a 10-m plant-free border
Avoid sequential planting because thrips can continue to emerge from the soil for 23
weeks after crop residues are ploughed and roto-tilled. Keep infected field areas fallow for
3-4 weeks to allow thrips to emerge from crop debris and disperse from the field.
During the first stage of plant production, use a net house structure or seedbed covered
with a netting of 40-mesh or higher to exclude thrips from seedlings prior to
transplanting. Use virus free transplants.
Several insecticide applications should be made at 5-day intervals to significantly reduce
an infestation.
Page
Symptoms
27
Transmission: Thrips
control measures
Applying lime or gypsum can provide calcium. Lime
corrects both low pH and low calcium levels, while
gypsum only affects calcium level.
Calcium chloride sprays applied to the foliage may help
prevent blossom-end rot on developing fruit. Calcium
chloride is applied at four pounds per 100 gallons/acre
four times on a weekly schedule, beginning when
symptoms first appear
Avoid excessive rates of ammonical nitrogen fertilizer.
Damage to the roots by deep cultivation should also be
avoided, especially after fruit set and in dry weather.
28
Page
Symptoms
Sunscald of Pepper
Page
Symptoms
29
References
Page
30
1. Vegetable diseases fact sheet, Edited by Tom Kalb. Photos by L.L. Black and E.L.Shannon. Published by AVRDC.
2. Vegetables 2014 final report. Plant Disease Diagnostic Center LSU/LSU AgCenter Department of Plant Pathology & Crop Physiology
Baton Rouge, Louisiana 70803.
3. Midwest Vegetable Production Guide for Commercial Growers Illinois University of Illinois Extension Indiana.
4. K.Pernezny, P.D. Roberts, J.F. Murphy and N.P. Goldberg(eds.), Compendium of pepper diseases. American Phytopathological Society
Press, St. Paul, MN.
Page
31
E. C. Stakman. 1959